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Grizzly bear 399 was fatally struck by a vehicle in Snake River Canyon, south of Jackson, Wyoming, last Tuesday.
The oldest known grizzly mother in history, the 28-year-old bear was often photographed with her cubs in Grand Teton National Park before she was killed by a car in October.
Bordered to the south by the Yellowstone and Lamar river basins, the Northern Range contains about 20 black bears for every 62 square miles.